When did the term "Princes in the Tower" come into usage, who invented it, and to whom did it refer? To the general public the term is synonymous with the boy King Edward V and his younger brother Richard, Duke of York, sons of Edward IV. Is this correct? Were those boys genuinely held against their will in the Tower? Would their mother, Elizabeth Widville, have released her young son Richard from sanctuary with her if she believed she would be putting his life in danger? By working exclusively with his own team of geneticists and exploring the mtDNA haplogroup of the living all-female-line collateral descendant of Edward V and his brother Richard, who has now been traced, John Ashdown-Hill answers these questions and more.